Oklahoma State University Center for Health Sciences
Oklahoma State University Center for Health Sciences

Anatomy and Vertebrate Paleontology

Anne Weil, Ph.D.Anne Weil, Ph.D.
Associate Professor of Anatomy




Education | Research Interests | Instructional Activities | Recent Publications





Ph.D. (Integrative Biology)
University of California, Berkeley
Berkeley, CA 

M.A. (Geological Sciences)
The University of Texas at Austin
Austin, TX

A.B. (English and American Literature and Language)
Harvard University
Cambridge, MA

Research Interests

I am a vertebrate paleontologist and paleobiologist. I study early mammals and recovery from extinction events.  I also am the senior scientist directing activities at the Homestead Site, a Jurassic locality in Oklahoma.

Early mammalian evolution, especially phylogeny and biogeography of multituberculate mammals
Although the Cenozoic is called “The Age of Mammals,” mammalian history began in the Mesozoic, and early mammals were quite diverse.  Multituberculates, sometimes called the “rodents of the Mesozoic,” lived on all the northern continents by the Late Cretaceous.  They are the most diverse and most commonly identified terrestrial vertebrates that survive the end-Cretaceous extinction and re-diversify afterward.  The best terrestrial record of this time interval is from the Western Interior of North America, where the mammalian fossil record is exceptional and there are age constraints on many local and regional faunas.  I am particularly interested in how biogeographic variation in the Late Cretaceous might have contributed to recovery of terrestrial ecosystems in the earliest Cenozoic.

Homestead Site, and Jurassic Morrison Formation biota from Oklahoma
First discovered in 2012, the Homestead Site has thus far produced fossils of Apatosaurus, Allosaurus, turtles, and crocodilians, as well as coprolites, snails, and ostracods.  Fossils are catalogued at the Sam Noble Museum in Norman.  I direct excavation by educational groups, mostly high school and college students, at the site, using their natural fascination with dinosaurs to teach about scientific questions, hypothesis tests, and methods, as well as about evolutionary themes. 

The first day of discovery at the site can be seen here: https://www.youtube.com/watch?v=Bsu1VmmR3z0; and a feature on 2015 activities here: http://scienceisok.com/digging-for-dinos/ .

Instructional Activities


  • Clinical Anatomy (team taught) – MS I
  • Anatomical enrichment activity for Ob/Gyn residents
  • Cornerstones in Vertebrate Paleontology – Graduate
  • Field Methods in Vertebrate Paleontology – Graduate
  • Advanced Vertebrate Paleontology (team-taught)  – Graduate
  • Statistics in Paleontology – Graduate
  • Vertebrate Osteology – Graduate


  • Gross and Developmental Anatomy – MS I
  • Biomedical Statistics – Graduate
  • Evolution and Development of the Mammalian Skull – Graduate

Recent Publications

Weil, A.  2014.  Mammalian evolution: A beast of the southern wild.  Nature 515:495-496.  doi:10.1038/nature13940

Williamson, T. E., Brusatte, S. L., Carr, T. D., Weil, A., and B. R. Standhardt.  2012. The phylogeny and evolution of Cretaceous-Paleogene metatherians: New cladistic analysis and description of new early Paleocene specimens from the Nacimiento Formation, New Mexico.  Journal of Systematic Palaeontology 10(4):625-651.  doi: 10.1080/14772019.2011.631592

Weil, A.  2011.  Mammalian evolution: A jaw-dropping ear. Nature 472:174-176.

Williamson, T. E. and A. Weil.  2011. A new early Paleocene (Puercan) hyopsodontid “condylarth” from New Mexico.  Acta Palaeontologica Polonica 56(2):247-255.  Available online 16 August 2010.  doi:10.4202/app.2009.0147

Williamson, T. E., A. Weil, and B. Standhardt. 2011. Cimolestids (Mammalia) from the early Paleocene (Puercan) of New Mexico. Journal of Vertebrate Paleontology 31(1):162-180.

Tanaka, K., Zelenitsky, D. K., Williamson, T. E., Weil, A. and F. Therrien.  2010.  Description of fossil eggshells from the Upper Cretaceous Fruitland Formation (upper Campanian), New Mexico. Historical Biology.  First publication 27 July 2010. doi:10.1080/08912963.2010.499171

Williamson, T. E., and Weil, A.  2008.  Stratigraphic distribution of sauropods in the Upper Cretaceous of the San Juan Basin, New Mexico, with comments on North America’s Cretaceous “sauropod hiatus.” Journal of Vertebrate Paleontology 28(4):1218-1223.

Williamson, T. E., and Weil, A.  2008.  Metatherian mammals from the Naashoibito Member, Kirtland Formation, San Juan Basin, New Mexico and their biochronologic and paleobiogeographic significance. Journal of Vertebrate Paleontology. 28(3):803-815.

Janis, C., and Weil, A.  2008.  Non-Eutherian mammals. pp. 7-18 in Janis, C., Gunnell, G., and Uhen, M. (eds.) Evolution of Tertiary Mammals of North America, Volume II.  Cambridge University Press, New York.

Weil, A. and Krause, D. W. 2008.  Multituberculata. pp. 19-38  in Janis, C., Gunnell, G., and Uhen, M. (eds.) Evolution of Tertiary Mammals of North America, Volume II. Cambridge University Press, New York.

Williamson, T. E., Nichols, D. J., and Weil, A.  2008.  Paleocene palynomorph assemblages from the Nacimiento Formation, San Juan Basin, New Mexico, and their biostratigraphic significance. New Mexico Geology 30(1):3-11.

Drea, C. and Weil, A.  2007, 2008.  External genital morphology of the Ringtailed Lemur (Lemur catta): females are naturally ‘masculinized’  Journal of Morphology, online publication Oct. 2007; print publication in April, 2008 269:451-463.



OSU-CHS on Facebook OSU-CHS on Twitter OSU-CHS on Foursquare OSU Medicine on YouTube Google+